NEW YORK -- Josh Wainwright caught 10 passes for 193 yards and two touchdowns, including a 24-yarder from Anders Hill in overtime, and Columbia defeated Pennsylvania 34-31 on Saturday for its best start since 1996.

The Lions (5-0, 2-0 Ivy) knocked off the defending Ivy League champions for the first time in 21 tries and snapped an 11-game home skid against the Quakers (2-3, 0-2). The Lions are assured of no worse than a .500 record after winning just five games in its last four seasons.

Penn appeared to be in control after Nick Miller's 25-yard interception return gave the Quakers a 21-7 lead in the third quarter. But the Lions rallied for three TDs in the fourth to take a 28-21 lead. Josh Bean started the Columbia scoring with a 1-yard plunge. Hill hit Wainwright for a 59-yard score to knot the score at 21 and then connected with Rory Schlageter from 7 yards out for the go-ahead score.

The Quakers sent the game to overtime on Will Fischer-Colbrie's 17-yard scoring strike to Justin Watson with 1:21 left. Penn got the ball first in OT, but settled for a 41-yard field goal. The Lions faced a third-and-9 when Hill hooked up with Wainwright for the game-winner.

Hill finished with 258 yards on 21-of-33 passing with three TDs and an interception.

Fischer-Colbrie completed 14 of 25 passes for 186 yards and a score. He threw two interceptions.

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