PHILADELPHIA -- Kyle Wilcox ran 20 yards for the game-winning touchdown in the fourth overtime as Penn topped Dartmouth 37-31 Saturday in the longest football game in Ivy League history.

Dartmouth had a chance to win in regulation, but David Park blocked Riley Lyons' 21-yard field goal as time expired for Penn (2-1, 1-0 Ivy). Dartmouth had another opportunity to end the game in the first overtime after stopping the Quakers on a fourth-and-1 from the Dartmouth 5. But Lyons then missed a 34-yard field goal that would have won the game for Dartmouth (1-2, 0-1).

After a 12-yard touchdown pass from Dalyn Williams to Kirby Schoenthaler in the second overtime, Billy Ragone ran for a 27-yard score on third-and-12 to tie the game for Penn and send it to a third overtime, where the teams traded field goals.

Wilcox then scored the game-winner following a missed 42-yard field goal from Lyons.